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Grind 1 tablespoon of the flaxseed with 1/4 cup water in a spice grinder until fairly smooth. Transfer to a small cup.
Repeat with remaining 1 tablespoon flaxseed and another 1/4 cup water. Alternatively, stir 2 tablespoons flaxseed meal into 1/2 cup water.
Meanwhile, place delicata squash in a single layer on a parchment paper-lined baking sheet. Bake alongside crust until soft and browned on the edges, about 15 minutes.
Combine leek, mushrooms and 2 tablespoons water in a large skillet over medium heat. Cook, stirring frequently, until browned and softened, about 12 minutes.
Stir in garlic and soy sauce, and cook for 1 minute. Transfer to a medium bowl.
Stir in quinoa, butternut squash purée, sage, vinegar, pepper, 1 tablespoon of the yeast and remaining flaxseed mixture.
Scrape into tart crust and level top. Sprinkle with remaining 1 tablespoon yeast.
Arrange squash slices on top, pressing them lightly into filling.
Bake until browned and set, about 45 minutes. Cool for 15 minutes before serving.